The wheel bug (Arilus cristatus) is one of the largest terrestrial true bugs in North America, being up to 1.5 inches, or 38 mm, in length; it is the only member of its genus. A characteristic structure is the wheel-shaped pronotal armor. They are predators upon soft-bodied insects such as caterpillars, Japanese beetles, etc., which they pierce with their beak to inject salivary fluids that dissolve soft tissue. Because most of their prey are pests, wheel bugs are considered beneficial insects, although they can inflict a painful bite if handled carelessly.
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